Frullania monocera

Epiphytic or lithophytic, brown to olive-green. Stems irregularly pinnately branched. Mid stem and branch lateral leaf lobes oblong-elliptic, 0.7–1.2 mm long, 0.55–1 mm wide, broadly obtuse to subtruncate at apex, not angled away from substrate when moist, imbricate; cells oblong at base, rounded-quadrate, irregular or circular elsewhere, 10–20 µm long and 5–15 µm wide at margin, 15–38 µm long and 13–20 µm wide near centre, 30–68 µm long and 20–45 µm wide toward base, with trigones throughout, particularly prominent at base, with intermediate thickenings most frequently from centre to margins. Lobules falcate with longest axis diverging from stem, 0.3–0.53 mm long, 0.2–0.5 mm wide, spinose at apex; cells 13–20 µm long, 6–15 µm wide, irregular, with prominent trigones and intermediate thickenings throughout. Stylus 3 cells long, 2 cells wide at base. Underleaves cuneate to obovate in outline, bifid to 2/5–1/2 of length, with a few spinose teeth along margins, 0.47–0.75 mm long, 0.4–0.83 mm wide, flat, with lobe apices pointing forwards; sinus acute; cells irregular, 10–38 µm long, 10–25 µm wide, smallest toward margins, with trigones and intermediate thickenings throughout. Perianth clavate-cylindric, 4-keeled, 1.8–2 mm long, 0.84–1 mm wide, truncate at apex, with a beak, covered with elongate tubercles; keels crenate.