Achrophyllum

Dioicous. Asexual reproduction occasionally by filamentous, often branched gemmae produced on on intramarginal cells of leaf surfaces. Mats on soil, rocks, logs or tree fern or tree trunks, often in running water. Stems creeping or erect, simple or with a few branches, with moderately dense rhizoids near base; pseudoparaphyllia present, foliose; central strand present. Leaves complanate, often most dense near stem apex, crisped when dry, those in lateral ranks oblong-ovate to spathulate, asymmetric at base, those in dorsal and ventral ranks elliptic or broadly ovate, symmetric, easily detached from stem; apex round, obtuse or rarely acute, without a hairpoint; costa present, extending to c. midleaf to ¾ of leaf length, usually shortly forked near midleaf; margins entire (not in Victoria) to dentate, plane, without a border; laminal cells hexagonal, becoming smaller toward margin and more elongate and rectangular near base, smooth. Seta smooth. Capsule pendent, symmetric, obolid, ellipsoid or ovoid, smooth when dry, with an annulus. Calyptra mitrate, smooth, with a fringed base. Operculum rostrate. Peristome double; endostome with segments as high as exostome teeth, with a basal membrane c. half of exostome height; cilia absent or rudimentary.