Tridontium
Dioicous or autoicous (not in Victoria). Asexual reproduction occasionally by clavate propagules in leaf axils. Turves, tufts or cushions on rock or soil, often in or near streams. Stems simple or branching irregularly, glabrous or with sparse rhizoids; central strand present or absent; sclerodermis present or absent; hyalodermis absent. Leaves ligulate-lanceolate, erect-spreading to recurved (not in Victoria) when moist, contorted and loosely appressed or with incurved apices when dry; apex round to bluntly acute; costa subpercurrent, with elongate adaxial superficial cells, without a differentiated adaxial epidermis, without an adaxial stereid band or adaxial stereid band weak, without a hydroid strand, with an abaxial stereid band, with a differentiated abaxial epidermis, with elongate abaxial superficial cells; margin entire, plane to erect toward apex, recurved in basal quarter, with c. 4 rows of enlarged cells forming an intramarginal border in basal third or half or sometimes without a border; laminal cells in apical half irregularly hexagonal or quadrate, smooth or uni-(not in Victoria) or pluripapillose (not in Victoria), unistratose throughout, with yellow or orange KOH reaction; basal laminal cells differentiated equidistantly from base from intramarginal border to costa, rhomboidal to rectangular, quadrate or shortly oblong at margin, longer, narrower and often coloured in intramarginal border. Acrocarpous. Capsule erect, straight, cupulate or urceolate to ovoid or cylindric (not in Victoria), exserted, operculate, with an annulus. Calyptra cucullate. Operculum apiculate or rostrate. Peristome absent (not in Victoria), rudimentary (not in Victoria), or of 16 teeth, cleft 2–3 times halfway or to base, straight.
Five species in New Guinea, south-east Australia, New Zealand, and Auckland and Macquarie Islands.
Tridontium has been placed in Grimmiaceae based on its resemblance to Scouleria (Zander 1993). However, phylogenies of DNA sequence data clearly place it within the Pottiaceae, closely related to Trichostomopsis and Didymodon (Cox et al. 2010; Jiménez et al. 2021). Originally conceived as a monotypic genus, two closely related species of Didymodon have been tranferred to Tridontium and another two species await formal description (Jiménez et al. 2021).
Cox, C.J.; Goffinet, B.; Wickett, N.J.; Boles, S.B.; Shaw, A.J. (2010). Moss diversity: A molecular phylogenetic analysis of genera. Phytotaxa 9: 175–195.
Jiménez, J.A.; Cano, M.J.; Guerra, J. (2021). A multilocus phylogeny of the moss genus Didymodon and allied genera (Pottiaceae): Generic delimitations and their implications for systematics. Journal of Systematics and Evolution.
Zander, R.H. (1993). Genera of the Pottiaceae: Mosses of harsh environments. Bulletin of the Buffalo Society of Natural Sciences 32: 1–378.