Dioicous. Asexual reproduction regularly by multicellar gemmae borne on costa adaxially. Tufts, mats or loose colonies on bark of trees or on twigs and small branches. Stems simple or sparingly branched, sparsely covered in rhizoids to densely tomentose near base; central strand absent; sclerodermis present; hyalodermis present. Leaves ovate to oblong-lanceolate, erect- or wide-spreading when moist, twisted and crisped when dry; apex shortly acuminate or acute; costa excurrent as a mucro, with short-rectangular superficial adaxial cells, with a differentiated adaxial epidermis, without an adaxial stereid band, with a hydroid strand, with an abaxial stereid band, without a differentiated abaxial epidermis, with elongate superficial abaxial cells; margin entire, plane or undulate, with a border of 3–4 rows of hyaline or yellow, smooth and more elongated cells, becoming intramarginal toward apex, with a row of smaller quadrate cells on the margin; laminal cells in apical half irregularly polygonal, rounded at angles, pluripapillose, KOH reaction red or yellow with red blotches or yellow-orange; basal laminal cells abruptly differentiated, rising higher marginally and fusing with border, rectangular or linear, smooth. Acrocarpous. Capsule erect, straight or curved, oblong-cylindric, exserted, operculate, with an annulus. Calyptra cucullate. Operculum conic. Peristome of 32 filaments, twisted anticlockwise, sometimes detaching with operculum.
