Sanionia
Autoicous. Asexual propagules absent. Tufts or mats on soil, logs or tree bases. Stems creeping or ascending to erect, irregularly or pinnately branched ± in one plane or rarely simple, with sparse rhizoids at or just below leaf insertions; paraphyllia absent; pseudoparaphyllia foliose; central strand present. Leaves lanceolate, circinate, falcate or almost straight, mostly falcate-secund when moist, not or scarcely altered when dry, strongly to weakly plicate, not or scarcely decurrent, branch leaves smaller than stem leaves often with a reduced alar group otherwise similar; apices acuminate to very long-acuminate; costa single, ending in acumen; margins denticulate or serrate (not in Victoria) throughout, or entire near base, plane near base, slightly recurved toward apex, without a border; laminal cells linear, shorter and broader at base, smooth, unistratose; alar cells clearly differentiated, inflated, quadrate to rectangular, hyaline, sharply delimited or not (not in Victoria), forming a transversely triangular or ±isodiametric (not in Victoria) group. Capsules erect to pendent, curved or straight, cylindric, with an annulus. Calyptra cucullate, smooth, glabrous. Opercula conic or conic-apiculate. Peristome double; exostome of 16 well developed entire teeth or reduced (not in Victoria); endostome of 16 well-developed alternating segments the same height as the exostome teeth or reduced (not in Victoria), with a high or low (not in Victoria) basal membrane; cilia well-developed, rudimentary (not in Victoria) or absent (not in Victoria).
Five species have been recognised within Sanionia in temperate and polar regions of both northern and southern hemispheres. However, four of these species likely evolved from S. uncinata based on phylogenetic analyses of chloroplast and nuclear DNA phylogenies and so in its current circumscription S. uncinata is composed of several lineages closer related to other species than they are to each other (Hedenäs 2012). The apparent lack of morphological distinctions (Hedenäs 2012) between these lineages makes this paraphyly hard to rectify by description of additional species currently included within S. uncinata
Hedenäs, L. (2012). Global phylogeography in Sanionia uncinata (Amblystegiaceae: Bryophyta). Botanical Journal of the Linnean Society 168: 19–42.