Brunonia

Perennial herbs with appressed hairs; hairs multicellular with terminal cell papillate. Leaves mostly basal, rosetted, villous in axils. Inflorescences head-like, consisting of several shortly pedunculate cymes, subtended by a long scape, surrounded by an involucre of bracts. Flowers sessile, erect, actinomorphic, bisexual, subtended by 4 membranous hyaline bracteoles; sepals 5, connate, subulate, plumose; petals 5, connate in lower half; stamens 5, opposite sepals, attached to petals; anthers connate, forming a tube, 2-celled, opening by longitudinal slits; ovary superior, 1-celled; ovule 1, attached basally on septum or ridge; style simple, entire with hollow indusium enclosing the more or less bifid tip. Fruit a nut enclosed in persistent calyx; seed without endosperm; testa membranous; embryo straight.