Cotula

Annual or perennial herbs. Leaves mostly cauline, alternate, entire to pinnatisect. Capitula cup-shaped or hemispherical, terminal or axillary, solitary, pedunculate; involucral bracts 2-seriate, subequal, herbaceous with scarious margins; receptacle flat, convex or conical, naked. Outer florets pedicellate, female, tubular, 1–many-seriate, rarely absent; corolla usually absent; style branches linear with papillose apices. Inner florets subsessile, bisexual or functionally male, tubular; corolla (3–)4-lobed, yellow; anthers obtuse or acute at base, with an ovate or lanceolate apical appendage; style with oblong branches, truncate, entire in sterile florets. Cypselas dorsally compressed, often winged, stipitate, with blunt-tipped hairs on surface; pappus absent.