Phascum
Autoicous or paroicous. Asexual propagules absent. Small scattered plants or turves on soil. Stem simple or often branched by innovation, with rhizoids in a basal tuft; central strand absent or present; sclerodermis absent; hyalodermis absent. Leaves becoming larger toward stem apex, oblong, elliptic, ovate, lanceolate or obovate, at least lower leaves spreading from capsule, otherwise erect when moist, twisted, incurved or scarcely altered when dry; apex obtuse, acute or acuminate, with or without a hairpoint; costa excurrent, with quadrate or rectangular adaxial superficial cells, with rectangular or elongate abaxial superficial cells, with an abaxial stereid band, with a hydroid strand, with a differentiated adaxial epidermis, with a differentiated abaxial epidermis; margin entire, revolute, recurved or plane, without a border; laminal cells in apical half quadrate, pentagonal, hexagonal or rhomboid, sometimes more elongate near apex, pluripapillose or rarely smooth, with yellow KOH reaction; basal laminal cells differentiated equidistant from base from margin to costa, rectangular, smooth. Acrocarpous. Capsules erect or pendent, ovoid to globose, with a blunt apiculus, immersed or emergent, cleistocarpous. Calyptra small, cucullate or occasionally mitrate, smooth, glabrous. Operculum absent. Peristome absent.
Around 20 species widespread through temperate regions; six species in Victoria.
Zander (1993) included Phascum in Tortula. Phascum is resolved among traditional Tortula species with operculate capsules and well-developed peristomes, including the type species, in phylogenies based on rps4 chloroplast DNA sequences (Werner et al. 2002), demonstrating the close relationship between these groups. However, in the same phylogenies, other genera retained as distinct by Zander (1993), such as Crossidium and Pterygoneuron, are also resolved among species that would belong to Tortula according to Zander (1993), suggesting that the circumscription of Tortula presented by Zander (1993) may also require change. Phylogenetic reconstruction of this group of mosses is still in its infancy with few species having been sampled and only one chloroplast region having been sequenced. Until Tortula and its close relatives are further studied in more detail, and the boundaries of Tortula are better clarified, Phascum is tentatively retained here.
Werner, O.; Ros, R.M.; Cano, M.J.; Guerra, J. (2002). Tortula and some related genera (Pottiaceae, Musci): phylogenetic relationships based on chloroplast rps4 sequences. *Plant Systematics and Evolution * 235: 197–207.
Zander, R.H. (1993). Genera of the Pottiaceae: Mosses of harsh environments. Bulletin of the Buffalo Society of Natural Sciences 32: 1–378.